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1、忍冬科忍冬科Caprifoliaceae中国植物志:13(12)/500(200+)FOC:5(66) /5(207)APG:42/890Largelytemperate/warmtemperateinthenorthernhemisphere,somemontanetropics(exp.Valerianoideae),butneithertheAntipodesnorthePacific.大洋洲太平洋五福花科锦带花亚科败酱亚科川续断亚科忍冬亚科北极花亚科刺续断亚科六道木属中国植物志接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核
2、木属Symphoricarpos鬼吹箫属Leycesteria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木属Zabelia糯米条属Abelia 双盾木属Dipelta锦带花属Weigela FOC接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核木属Symphoricarpos鬼吹箫属Leycesteria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木属Zabelia糯米条属Abelia 双盾木属Dipelta双六道木属Diabelia 锦带花属Weigela 五福花科Ad
3、oxaceae北极花科Linnaeaceae锦带花科Diervillaceae忍冬科Caprifoliaceae双六道木属Diabelia 忍冬科Cymeswithapairofbractsand2pairsofbracteoles,occasionallyaccrescentinfruit,rarelyabsent.anthersfree,2-celled,Ovaryinferior,Fruitaberry,adrupewith2-5pyrenes,oraleatheryachene.Seeds1tomany;五福花科Inflorescencesterminalpaniclesorumbel
4、late,spicate,orcapitatecymes.anthersfree,2-celled,Ovarysemi-inferiortoinferior,Fruitdrupes;seeds1or3-5.锦带花科:Ovaryinferior,withagloboseglandononesideofupperpart,Fruita2-valveddehiscentcapsule北极花科:bractsleaflikeorreducedandlocatedatbaseofovaryandforminganepicalyx.1.花序由聚散合成伞形、伞房式或圆锥式;柱头常2-3裂;花冠不具蜜腺;花药内
5、向或外向五福花科1.花序非上述情况,若为圆锥花序则花柱细长;柱头大多为头状,很少分裂;花冠具蜜腺;花药内向。2.蒴果锦带花科2.核果。3.聚伞花序有一对苞片和两对小苞片,稀缺;种子1到多数忍冬科3.苞片叶状,着生在子房基部形成萼状总苞;种子1-颗北极花科五福花科Adoxaceae1.Leavessimple;ovary1-loculed;drupes1-seeded;shrubsorsmalltreesViburnum 1.Leavescompound;ovary3-5-loculed;drupesusually3-5-seeded;shrubsorsmalltreesSambucus五福花科
6、Adoxaceae-荚蒾属Viburnum单叶核果1种子花药内向叶柄及花冠筒被星状毛,花芳香V. dilatatum荚蒾V. macrocephalum绣球荚蒾V. odoratissimum珊瑚树叶革职不育花五福花科Adoxaceae-接骨木属SambucusS. williamsii接骨木奇数羽状复叶花药外向核果浆果状Adoxa moschatellina五福花草本,复叶花雄蕊2轮,内轮退化,外轮分裂为2半蕊花药外向中国植物志接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核木属Symphoricarpos鬼吹箫属Leyceste
7、ria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木属Zabelia糯米条属Abelia 双盾木属Dipelta锦带花属Weigela FOC接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核木属Symphoricarpos鬼吹箫属Leycesteria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木属Zabelia糯米条属Abelia 双盾木属Dipelta双六道木属Diabelia 锦带花属Weigela 五福花科Adoxaceae北极花科Linnaeaceae锦带花科Di
8、ervillaceae忍冬科Caprifoliaceae双六道木属Diabelia 忍冬科Caprifoliaceae1.Flowerssingleandoppositeinaspike2.Ovaries5,7,or8-locular,allloculeswithmanyovules. Leycesteria鬼吹箫属2.Ovaries4-locular,2loculeswith1fertileovule,other2loculeswithmanysterileovules. Symphoricarpos毛核木属1.Flowerspaired(rarelysingleandthenpeduncu
9、late;rarelyinthreesandthenbracteolesfusedintoacupule)orinwhorlsof63.Flowerspaired(rarelysingleandthenpedunculate;rarelyinthreesandthenbracteolesfusedintoacupule). Lonicera忍冬属3.Flowersinwhorlsof64.Flowers6inasinglewhorl,with2pairsoftightinvolucralbractsformingacapitulum;capitulapaniculate;shrubswitht
10、riplinervedleaves.Heptacodium七子花属小苞片连合成杯状而包被萼筒两对紧密缩合的苞片组成头状的圆锥花序忍冬科Caprifoliaceae-毛核木属SymphoricarposS. sinensis毛核木灌木,花单生于短小、钻形苞片的腋内,形成顶生穗状花序果顶端有一小喙,蓝黑色,具白霜忍冬科Caprifoliaceae1.Flowerssingleandoppositeinaspike2.Ovaries5,7,or8-locular,allloculeswithmanyovules. Leycesteria鬼吹箫属2.Ovaries4-locular,2loculesw
11、ith1fertileovule,other2loculeswithmanysterileovules. Symphoricarpos毛核木属1.Flowerspaired(rarelysingleandthenpedunculate;rarelyinthreesandthenbracteolesfusedintoacupule)orinwhorlsof63.Flowerspaired(rarelysingleandthenpedunculate;rarelyinthreesandthenbracteolesfusedintoacupule). Lonicera忍冬属3.Flowersinwh
12、orlsof64.Flowers6inasinglewhorl,with2pairsoftightinvolucralbractsformingacapitulum;capitulapaniculate;shrubswithtriplinervedleaves.Heptacodium七子花属小苞片连合成杯状而包被萼筒两对紧密缩合的苞片组成头状的圆锥花序忍冬科Caprifoliaceae-七子花属HeptacodiumOnespeciesH. Miconioides 七子花叶三出脉宿存花萼被刺刚毛花七朵,花冠稍二唇1.Flowerssingleandoppositeinaspike2.Ovari
13、es5,7,or8-locular,allloculeswithmanyovules. Leycesteria鬼吹箫属2.Ovaries4-locular,2loculeswith1fertileovule,other2loculeswithmanysterileovules.Symphoricarpos毛核木属1.Flowerspaired(rarelysingleandthenpedunculate;rarelyinthreesandthenbracteolesfusedintoacupule)orinwhorlsof63.Flowerspaired(rarelysingleandthen
14、pedunculate;rarelyinthreesandthenbracteolesfusedintoacupule). Lonicera忍冬属3.Flowersinwhorlsof64.Flowers6inasinglewhorl,with2pairsoftightinvolucralbractsformingacapitulum;capitulapaniculate;shrubswithtriplinervedleaves.Heptacodium七子花属忍冬科Caprifoliaceae小苞片连合成杯状而包被萼筒两对紧密缩合的苞片组成头状的圆锥花序4.Flowers6inasinglew
15、horl,with2pairsoftightinvolucralbractsformingacapitulum;capitulapaniculate;shrubswithtriplinervedleaves.Heptacodium七子花属4.Inflorescencedifferent,ifflowersinacapitulumthenplantaclimberwithperfoliateinvolucralbractsandbilabiateflowersin1-4whorls5.Herbs;leavespinnatifid,otherwiseperfoliate Triosteum莛子藨属
16、5.Shrubsorclimbers;leavesentire,notperfoliate(involucralbractssometimesperfoliate).6.Shrubs;flowersfunnelform Leycesteria鬼吹箫属6.Climbers;flowersbilabiate Lonicera忍冬属抱茎的总苞二唇的羽裂的漏斗状的忍冬科Caprifoliaceae-鬼吹箫属Leycesteria示蜜腺囊穗状花序每节对生子房7-8室,每室胚珠多数L. gracilis纤细鬼吹箫忍冬科Caprifoliaceae-鬼吹箫属LeycesteriaL. formosa鬼吹箫花
17、每节6朵,苞片叶状,每轮6枚花冠漏斗状果具宿存萼齿忍冬科Caprifoliaceae1.Flowerssingleandoppositeinaspike2.Ovaries5,7,or8-locular,allloculeswithmanyovules. Leycesteria鬼吹箫属2.Ovaries4-locular,2loculeswith1fertileovule,other2loculeswithmanysterileovules. Symphoricarpos毛核木属1.Flowerspaired(rarelysingleandthenpedunculate;rarelyinthre
18、esandthenbracteolesfusedintoacupule)orinwhorlsof63.Flowerspaired(rarelysingleandthenpedunculate;rarelyinthreesandthenbracteolesfusedintoacupule). Lonicera忍冬属3.Flowersinwhorlsof64.Flowers6inasinglewhorl,with2pairsoftightinvolucralbractsformingacapitulum;capitulapaniculate;shrubswithtriplinervedleaves
19、.Heptacodium七子花属小苞片连合成杯状而包被萼筒两对紧密缩合的苞片组成头状的圆锥花序4.Flowers6inasinglewhorl,with2pairsoftightinvolucralbractsformingacapitulum;capitulapaniculate;shrubswithtriplinervedleaves.Heptacodium七子花属4.Inflorescencedifferent,ifflowersinacapitulumthenplantaclimberwithperfoliateinvolucralbractsandbilabiateflowersin
20、1-4whorls5.Herbs;leavespinnatifid,otherwiseperfoliate Triosteum莛子藨属5.Shrubsorclimbers;leavesentire,notperfoliate(involucralbractssometimesperfoliate).6.Shrubs;flowersfunnelform Leycesteria鬼吹箫属6.Climbers;flowersbilabiate Lonicera忍冬属抱茎的总苞二唇的羽裂的漏斗状的L. japonica忍冬忍冬科Caprifoliaceae-忍冬属Lonicera花二唇形苞片忍冬科Cap
21、rifoliaceae-忍冬属LoniceraL. tragophylla盘叶忍冬花序下1-2对叶连合成盘状3朵花组成聚伞花序密集成头状花序4.Flowers6inasinglewhorl,with2pairsoftightinvolucralbractsformingacapitulum;capitulapaniculate;shrubswithtriplinervedleaves.Heptacodium七子花属4.Inflorescencedifferent,ifflowersinacapitulumthenplantaclimberwithperfoliateinvolucralbrac
22、tsandbilabiateflowersin1-4whorls5.Herbs;leavespinnatifid,otherwiseperfoliate Triosteum莛子藨属5.Shrubsorclimbers;leavesentire,notperfoliate(involucralbractssometimesperfoliate).6.Shrubs;flowersfunnelform Leycesteria鬼吹箫属6.Climbers;flowersbilabiate Lonicera忍冬属抱茎的总苞二唇的羽裂的漏斗状的忍冬科Caprifoliaceae-莛子藨属Triosteum
23、 T. pinnatifidum莛子藨草本,叶羽状深裂果具宿存萼齿3多花组成的聚伞花序对生中国植物志接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核木属Symphoricarpos鬼吹箫属Leycesteria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木属Zabelia糯米条属Abelia 双盾木属Dipelta锦带花属Weigela FOC接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核木属Symphoricarpos鬼吹箫属Leyces
24、teria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木属Zabelia糯米条属Abelia 双盾木属Dipelta双六道木属Diabelia 锦带花属Weigela 五福花科Adoxaceae北极花科Linnaeaceae锦带花科Diervillaceae忍冬科Caprifoliaceae双六道木属Diabelia 北极花科Linnaeaceae1.Petiolesofoppositeleafpairsdilatedandconnateatbase,enclosingaxillarybuds;corollahypocrateriform,withacylindr
25、icaltubeand4or5spreadinglobes;flowersin3-floweredcymesorpaired;stigmasgreenandmucilaginousZabelia六道木属1.Petiolesofoppositeleafpairsnotdilated,axillarybudsexposed;corollafunnelformorbilabiatewith5spreadinglobes;flowerspairedorsingle;stigmaswhiteandnotmucilaginous.2.Twobractsaccrescentandbecomingwingli
26、keinfruit;ovary4-locularwith2fertileloculesDipelta双盾木属2.Bractsnotwinglikeinfruit;ovary3-or4-locularwith1or2fertilelocules.扩大合生高脚碟形圆柱形的分泌粘液漏斗形北极花科Linnaeaceae-六道木属ZabeliaTheZ. bifloraspeciescomplex花冠高脚碟形,柱头绿色叶柄基部膨大合生花被裂片4宿存花萼疏被纤毛TheZabelia bifloraspeciescomplexwasrevisedbyNakai(J.Jap.Bot.13(8):1-12.19
27、37)whodistinguished11species.Wecurrentlyrecognizetwospecies(Zabelia biflora(Turczaninow)Makino六道木andZabelia dielsii(Graebner)Makino南方六道木)inChina.Mostofthedifferencesbetweentaxawerebasedonthelengthofthepeduncles,bracts,andbracteoles,whicharehighlyvariable.ClassificationofZabelia bifloraiscomplexandne
28、edstobeinvestigated.北极花科Linnaeaceae-六道木属ZabeliaZ. triflora醉鱼草状六道木宿存萼片密被毛花被裂片5北极花科Linnaeaceae1.Petiolesofoppositeleafpairsdilatedandconnateatbase;corollahypocrateriform,withacylindricaltubeand4or5spreadinglobes;flowersin3-floweredcymesorpaired;stigmasgreenandmucilaginousZabelia六道木属1.Petiolesofopposit
29、eleafpairsnotdilated;corollafunnelformorbilabiatewith5spreadinglobes;flowerspairedorsingle;stigmaswhiteandnotmucilaginous.2.Twobractsaccrescentandbecomingwinglikeinfruit;ovary4-locularwith2fertileloculesDipelta双盾木属2.Bractsnotwinglikeinfruit;ovary3-or4-locularwith1or2fertilelocules.扩大合生高脚碟形圆柱形的分泌粘液漏斗
30、形北极花科Linnaeaceae-双盾木属DipeltaD. floribunda双盾木小苞片盾状,膜质,宿存而增大成翅状花冠稍二唇形北极花科Linnaeaceae1.Petiolesofoppositeleafpairsdilatedandconnateatbase;corollahypocrateriform,withacylindricaltubeand4or5spreadinglobes;flowersin3-floweredcymesorpaired;stigmasgreenandmucilaginousZabelia六道木属1.Petiolesofoppositeleafpairs
31、notdilated;corollafunnelformorbilabiatewith5spreadinglobes;flowerspairedorsingle;stigmaswhiteandnotmucilaginous.2.Twobractsaccrescentandbecomingwinglikeinfruit;ovary4-locularwith2fertileloculesDipelta双盾木属2.Bractsnotwinglikeinfruit;ovary3-or4-locularwith1or2fertilelocules.扩大合生高脚碟形圆柱形的分泌粘液漏斗形3.Evergre
32、en,creepingsubshrubs;twobractsshieldlikeandcoveredwithstickyglandularhairs;inflorescencereducedto2singleflowersatapicesofbranchlets(thosecanlooklikepairedflowersbuthavemorebractsandinternodesseparatingthem)Linnaea北极花属3.Erectshrubs;bractsandinflorescencenotasabove.4.Bractsandbracteolescoveredwithbris
33、tles,accrescentandbecomingspongy,woodyandfusedtoovariesinfruit.Kolkwitzia蝟实属4.Bractsandbracteoleswithoutbristles,smallandnon-accrescentinfruit.5.Flowersterminalandpaired(flowersopeningsimultaneously),sometimes1-3(-8)duetosupernumeraryflowersaxillarytobracteoles(moreoftensoonrepeat-bloomlongshoots),s
34、pringfloweringDiabelia双六道木属5.Flowersaxillary,paniculate,singleorpaired(flowersopeningconsecutively),summerorautumnfloweringAbelia糯米条属北极花科Linnaeaceae-北极花属LinnaeaL. borealis北极花Onespecies苞片盾状,密被粘性腺毛果实下垂匍匐亚灌木3.Evergreen,creepingsubshrubs;twobractsshieldlikeandcoveredwithstickyglandularhairs;inflorescenc
35、ereducedto2singleflowersatapicesofbranchlets(thosecanlooklikepairedflowersbuthavemorebractsandinternodesseparatingthem)Linnaea北极花属3.Erectshrubs;bractsandinflorescencenotasabove.4.Bractsandbracteolescoveredwithbristles,accrescentandbecomingspongy,woodyandfusedtoovariesinfruit.Kolkwitzia蝟实属4.Bractsand
36、bracteoleswithoutbristles,smallandnon-accrescentinfruit.5.Flowersterminalandpaired(flowersopeningsimultaneously),sometimes1-3(-8)duetosupernumeraryflowersaxillarytobracteoles(moreoftensoonrepeat-bloomlongshoots),springfloweringDiabelia双六道木属5.Flowersaxillary,paniculate,singleorpaired(flowersopeningco
37、nsecutively),summerorautumnfloweringAbelia糯米条属K. amabilis蝟实北极花科Linnaeaceae-蝟实属KolkwitziaOnespecies花冠5裂,不等大苞片小苞片密被刺刚毛,宿存萼齿宿存,2朵花组成的聚伞花序伞房状3.Evergreen,creepingsubshrubs;twobractsshieldlikeandcoveredwithstickyglandularhairs;inflorescencereducedto2singleflowersatapicesofbranchlets(thosecanlooklikepaired
38、flowersbuthavemorebractsandinternodesseparatingthem)Linnaea北极花属3.Erectshrubs;bractsandinflorescencenotasabove.4.Bractsandbracteolescoveredwithbristles,accrescentandbecomingspongy,woodyandfusedtoovariesinfruit.Kolkwitzia蝟实属4.Bractsandbracteoleswithoutbristles,smallandnon-accrescentinfruit.5.Flowerste
39、rminalandpaired(flowersopeningsimultaneously),springfloweringDiabelia双六道木属5.Flowersaxillary,paniculate,singleorpaired(flowersopeningconsecutively),summerorautumnfloweringAbelia糯米条属圆锥花序北极花科Linnaeaceae-糯米条属Abelia A. chinensis糯米条A. grandiflora大花糯米条A. chinensis A. uniflora蓪梗花花成对腋生,雄蕊明显伸出花冠筒花单生或双生,雄蕊稍伸出花
40、冠筒中国植物志接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核木属Symphoricarpos鬼吹箫属Leycesteria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木属Zabelia糯米条属Abelia 双盾木属Dipelta锦带花属Weigela FOC接骨木属Sambucus荚蒾属Viburnum 莛子藨属Triosteum 七子花属Heptacodium毛核木属Symphoricarpos鬼吹箫属Leycesteria忍冬属Lonicera北极花属Linnaea蝟实属Kolkwitzia六道木
41、属Zabelia糯米条属Abelia 双盾木属Dipelta双六道木属Diabelia 锦带花属Weigela 五福花科Adoxaceae北极花科Linnaeaceae锦带花科Diervillaceae忍冬科Caprifoliaceae双六道木属Diabelia 锦带花科Diervillaceae-锦带花属Weigela Ovaryinferior,withagloboseglandononesideofupperpart,Fruita2-valveddehiscentcapsule.锦带花科Diervillaceae-锦带花属Weigela W. florida锦带花W. japonica半边月花萼裂至中部,裂齿披针形花萼裂至基部,裂齿条形